Koko the famous low land gorilla who learned sign language died peacefully in her sleep at the age of 46.

“Her impact has been profound and what she has taught us about the emotional capacity of gorillas and their cognitive abilities will continue to shape the world,” the Gorilla Foundation said.

Source: https://www.theguardian.com/world/2018/jun/21/koko-gorilla-sign-language-dead-aged-46